一、教育经历

1990.09–1994.06，四川大学生物系，获学士学位；

1996.09–1999.06，江西农业大学林学院，获硕士学位；

2000.09-2003.07，中国科学院昆明植物研究所，获博士学位；

2003.11–2006.12，中国科学院植物研究所，博士后

2014.09–2015.09，美国佛罗里达大学自然历史博物馆，访问学者

二、工作经历

1994.07–2022.12，江西农业大学农学院，助教、讲师、副教授、教授

2022.12–至今 广西师范大学生命科学学院，教授、博导；珍稀濒危动植物生态与环境保护教育部重点实验室，副主任

三、人才称号/学术头衔/社会兼职

广西高层次人才（C层次），江西省全省模范教师、江西省主要学科学术与技术带头人（领军人才）、江西省青年科学家、江西省五四青年奖章获得者；入选江西省百千万人才工程；广西植物学会常务理事，江西植物学会常务理事；江西省自然科学一等奖获得者（排名第一）。

四、教学情况

本科生课程：《植物学》

研究生课程：《分子生态学》《植物分类学原理与方法》

五、科研项目

1．主持国家自然科学基金“两个长柄水青冈与亮叶水青冈种间杂交带的比较研究：生殖隔离位点与适应性性状的解析”（2024–2027，批准号：32360064）

2．主持完成国家自然科学基金“杜鹃花科吊钟花属多倍化和系统发育研究”（2020–2023，批准号：31960049）

3.主持完成国家自然科学基金“世界水青冈属物种界定、系统发育和生物地理研究”（2018–2021，批准号：31770236）

4.主持完成国家自然科学基金“中国亚热带喜冷植物裂叶星果草谱系地理学研究”（2016–2019，批准号：31560064）

5.主持完成国家自然科学基金“江西森林植物多物种遗传多样性分布格局研究”（2012–2015，批准号：31160043）

6.主持完成国家自然科学基金“中国几种水青冈分子谱系地理学研究”（2008–2010，30760016）

7.主持完成国家自然科学基金“三种濒危五针松保护遗传学研究”（2005–2007，批准号30460020）

8.主持广西科技基地和人才专项—高层次创新人才和团队培养“杜鹃花科吊钟花属植物资源产业化利用关键技术研究（2025–2027，批准号：2024AC43009）

9.主持完成江西省主要学科学术与技术带头人（领军人才）培养计划项目：杜鹃花科吊钟花属多倍化与自然杂交及其引种驯化研究（2021–2023，批准号：20213BCJ22006）

10.主持完成江西省教育厅“落地计划”项目“华木莲繁育系统和小尺度遗传结构研究”（2013–2015，批准号：KJLD12062）

11.主持完成江西省青年科学家（井冈之星）培养对象计划项目（2009–2011，2008DQ01500）

12.主持完成系统与进化植物学国家重点实验室开放课题“濒危植物华木莲繁育系统和遗传多样性研究”（2010–2011）

13.主持完成国家科技支撑计划“流域生物多样性保护及药用资源开发利用技术研究与示范”（科技部2012BAC11B02）子课题“流域生物多样性保护及药用资源开发利用技术研究与示范”

15.参加国家自然科学基金“中国亚热带常绿阔叶林3个优势种的比较亲缘地理学研究”（2012–2015，批准号：31160082）

六、代表性论著  

1.Hua Liang#, Cheng Zhou#, Wan Hu#, Sujuan Wei, Dengmei Fan, Shuang Tian*, Zhiyong Zhang*. 2025. Cytotype distribution and chloroplast phylogeography of Enkianthus chinensis complex, a mixed-ploidy species complex in subtropical China and adjacent areas. Perspectives in Plant Ecology, Evolution, and Systematics, https://doi.org/10.1016/j.ppees.2025.125909.

2.Rizal M. Suhardi, Li-Ping Ju, Tsai-Wen Hsu, Tze-Ying Chen, Huei-Chuan Shih, Zhi-Yong Zhang*, Ya-Zhu Ko*, Meng-Shin Shiao*, Yu-Chung Chiang*. 2025. Unraveling historical genetic divergence and gene flow patterns between island (Taiwan) and mainland (China) of Fagus hayatae. Frontiers in Plant Science, 16:1628728. doi: 10.3389/fpls.2025.1628728

3.Shuilian Peng, Zhijun Zhai, Wan Hu, Hua Liang, Yi Yang, Yixuan Kou, Meixia Wang, Shanmei Cheng, Zhiyong Zhang*, Dengmei Fan*. 2025. Complete chloroplast genome of Enkianthus Lour. (Ericaceae): Comparative analysis, phylogenetic relationships, divergence history, and adaptive evolution. Ecology and Evolution, 15:e72129. https://doi.org/10.1002/ece3.72129

4.Gulbar Yisilam,  Kenneth M. Cameron,  Zhi-Yong Zhang,  En-Ting Zheng,  Chuan-Ning Li,  Zhen-Zhou Chu,  Ying Su,  Jia-Lei Li,  Yu-Wei Wang,  Jin Li,  Pan Li,  Xin-Min Tian. 2025. New insights into the phylogeny and biogeography of Ggoji berries (Lycium, Solanaceae) inferred from plastid data. Journal of Biogeography,  https://doi.org/10.1111/jbi.15163

5.Gulbar Yisilam, Enting Zheng, Chuanning Li, Zhiyong Zhang, Ying Su, Zhenzhou Chu, Pan Li, Xinmin Tian. 2025. The chromosome-scale genome of Black Wolf berry (Lycium ruthenicum) provides useful genomic resources for identifying genes related to anthocyanin biosynthesis and disease resistance. Plant Diversity, 47:201-213. https://doi.org/10.1016/j.pld.2025.01.001.

6.Wan Hu, Qi Qiu, Hua Liang, Yang Liu, Yi Yang, Shudong Zhang, Dengmei Fan*, Zhiyong Zhang*. 2025. Speciation and hybridization of Enkianthus quinqueflorus and E. serrulatus (Ericaceae) across a tropical-subtropical transitional zone in south China. Botanical Journal of the Linnean Society, 209(2): 173-185. https://doi.org/10.1093/botlinnean/boaf013. 

7.Yi-Xuan Kou, Mi-Li Liu, Jordi Lopez-Pujol, Qi-Jing Zhang, Zhi-Yong Zhang, Zhong-Hu Li*. 2024. Contrasting demographic history and mutational load in three threatened whitebark pines (Pinus subsect. Gerardianae): implications for conservation. The Plant Journal. doi: 10.1111/tpj.16965.

8.Yanfang Lin, Yingying Xiang, Sujian Wei, Qiwei Zhang, Yanhua Liu, Zhiyong Zhang, Shaoqing Tang*. 2024. Genetic diversity and population structure of an insectpollinated and bird-dispersed dioecious tree Magnolia kwangsiensis in a fragmented karst forest landscape. Ecology and Evolution, 14:e70094.

9.Haimiao Wang, Li Huang, Pan Yang, Xianghua Zeng, Yuli Huang, Wenting Yuan, Yixuan Kou, Zhiyong Zhang*. 2024. Short-term soil waterlogging improves cotton tolerance to high temperature by triggering antioxidant defense system in cotton seedlings. Journal of Agronomy and Crop Science. 210:e12713.

10.Wei-Li Liu, Dan-Qi Li, Yi-Ping Zou, Ming-Zhuo Hao, Zhi-Yong Zhang, Deng-Mei Fan, Yi Yang. 2024. Hybrid origin and status of the endangered Ilex sanqingshanensis revealed by molecular and morphological evidence. Plant Systematics and Evolution. 310:16.

11. Cheng Zhou#, Hua Liang#, Wan Hu, Yi Yang, Xinmin Tian, Chie Tsutsumi, Dengmei Fan*, Zhiyong Zhang*. 2024. Molecular phylogeny of Enkianthus Lour. based on chloroplast and nuclear DNA sequences with an emphasis on the origin of polyploid species. Scientia Horticulturae, 328: 112960.

12.Danqi Li#, Lu Jiang#, Wei He, Dengmei Fan, Shanmei Cheng, Yi Yang,Meixia Wang, Shaoqing Tang, Yixuan Kou*, Zhiyong Zhang*. 2024. Allopatric speciation and secondary sympatry of Fagus longipetiolata and F. lucida (Fagaceae) in subtropical China. Botanical Journal of the Linnean Society. doi:10.1093/botlinnean/boad077.

13.Yixuan Kou, Dengmei Fan, Shanmei Cheng, Yi Yang, Meixia Wang, Yujin Wang*, Zhiyong Zhang*. 2023. Peripatric speciation within Torreya fargesii (Taxaceae) in the Hengduan Mountains inferred from multi-loci phylogeography. BMC Ecology and Evolution,  23:74.

14. Danqi Li#, Lu Jiang#, Hua Liang, Dahai Zhu, Dengmei Fan, Yixuan Kou, Yi Yang*, Zhiyong Zhang*. 2023. Resolving a nearly 90-year-old enigma: the rare Fagus chienii is conspecific with F. hayatae based on molecular and morphological evidence. Plant Diversity,45: 544–551.

15.Yi Yang, Lei Jiang, En-De Liu, Wei-Li Liu, Li Chen,Yi-Xuan Kou, Deng-Mei Fan, Shan-Mei Cheng, Zhi-Yong Zhang* and Hua Peng*. 2023. Time to update the sectional classification of Ilex (Aquifoliaceae): new insights from Ilex phylogeny, morphology and distribution. Journal of Systematics and Evolution.61: 1036–1046.

16.Hua Liang, Lu Jiang, Danqi Li, Yi Yang, Dengmei Fan*, Zhiyong Zhang*. 2022. A new synonym of Enkianthus (Ericaceae) in East Asia based on morphological and molecular evidence. PhytoKey, 214: 61–74.

17.Lu Jiang, Qin Bao, Wei He, Dengmei Fan, ShanmeiCheng, Jordi López-Pujol, Myong Gi Chung, Shota Sakaguchi, Arturo Sánchez González, Aysun Gedik, Dezhu Li, Yixuan Kou*, and Zhiyong Zhang*. 2022. Phylogeny and biogeographyof Fagus L. (Fagaceae) based on 28 nuclear single/low-copy loci. Journal of Systematics and Evolution. 60(4): 759–772.

18.Dengmei Fan, Shuqing Lei, Hua Liang, Qi Yao, YixuanKou*, Shanmei Cheng, Yi Yang, Yingxiong Qiu, Zhiyong Zhang*. 2022. More opportunities more species: Pleistocene differentiation and northward expansion of an evergreenbroadleaved tree species Machilus thunbergii Sieb. et Zucc. (Lauraceae)in southeast China. BMC Plant Biology, 22: 35.

19.Shanmei Cheng#, Weidong Zeng#, Jing Wang, Lei Liu, Hua Liang, Yixuan Kou, Hengchang Wang, Dengmei Fan*, ZhiyongZhang*. 2021. Species delimitation of Asteropyrum (Ranunculaceae) based on morphological, molecular, and ecological variation. Frontiers in Plant Science,12:681864.

20.Yixuan Kou, Li Zhang, Dengmei Fan, Dezhu Li,Richard G. J. Hodel, Zhiyong Zhang*. 2020. Evolutionary history of a relict conifer Pseudotaxus chienii (Taxaceae) during thelate Neogene. Annals of Botany, 125: 105-117.

21.Shuang Tian#, Yixuan Kou#, Zhirong Zhang#, Lin Yuan, Derong Li, Jordi López-Pujol, DengmeiFan*, ZhiyongZhang*. 2018. Phylogeography of Eomecon chionanthain subtropical China: the dual roles of the Nanling Mountains as a glacialrefugium and a dispersal corridor. BMC Evolutionary Biology, 18: 20.

22.Dengmei Fan#, Zhixia Sun#, Bo Li, Yixuan Kou, Richard G. J. Hodel, Zhirong Jin, ZhiyongZhang*. 2017.Dispersal corridors for plant species in the Poyang Lake Basin of southeast China identified by integration of phylogeographic and geospatial data. Ecology and Evolution, 7(14): 5140-5148.

23.Yi-Xuan Kou, Kun Xiao, Xiao-Rong Lai, Yu-Jin Wang, Zhi-Yong Zhang*. 2017. Natural hybridization between Torreya jackii and T.grandis (Taxaceae) in southeast China. Journal of Systematics and Evolution, 55: 25-33.

24.Dengmei Fan#, Wan Hu#, Bo Li, Ashley B. Morris,Min Zheng, Douglas E. Soltis, Pamela S. Soltis, Zhiyong Zhang*. 2016. Idiosyncratic responses of evergreen broad-leaved forest constituents in China to the late Quaternary climate changes. Scientific Reports, 6: 31044.

25.Yixuan Kou#, Shanmei Cheng#, Shuang Tian#, Bo Li, Dengmei Fan, Yongjun Chen, Douglas E. Soltis, Pamela S. Soltis, Zhiyong Zhang*.2016. The antiquity of Cyclocarya paliurus (Juglandaceae) provides new insights into the evolution of relict plants insubtropical China since the late early-Miocene. Journal of Biogeography,43: 351-360.

26.Shuang Tian#, Shuqing Lei#, Wan Hu#, Lingli Deng, Qinglin Meng, Bo Li, Douglas E. Soltis, Pamela S. Soltis, Dengmei Fan*, Zhi-Yong Zhang*. 2015.Repeated range expansions andinter-/postglacial recolonization routes of Sargentodoxa cuneata (Oliv.) Rehd. et Wils. (Lardizabalaceae) in subtropical China. Molecular Phylogenetics and Evolution, 85: 238-246.

27.胡菀, 罗意, 阳亿, 张志勇, 范邓妹*. 2014. 野生桂花的遗传多样性和遗传结构研究. 园艺学报.41(7): 1427-1435.

28.熊敏, 田双, 范邓妹, 张志勇*. 2014. 华木莲群体遗传结构和保护单元研究. 生物多样性. 22(4): 476-484.

29.Zhi-Yong Zhang*, Rong Wu, Qun Wang, Zhi-Rong Zhang, Deng-Mei Fan, Jordi López-Pujol, De-Zhu Li*.2013. Comparative phylogeography of two sympatric beeches in subtropical China: Species-specific geographic mosaic of lineages. Ecology and Evolution,3(13): 4461-4472.

30.Ming Lei, Qun Wang, Zhen-Jian Wu, JordiLópez-Pujol, De-Zhu Li*, and Zhi-YongZhang*. 2012. Molecular phylogeography of Fagus engleriana (Fagaceae) in subtropical China: limited admixture among multiple refugia. Tree Genetics andGenomes, 8: 1203-1212.

31.Shi-Quan Guo, Min Xiong, Zhi-RongZhang, De-Zhu Li & Zhi-Yong Zhang*. 2011. Molecular phylogenetic reconstruction of Osmanthus Lour. (Oleaceae) and related genera based on three chloroplast intergenic spacers. Plant Systematic and Evolution 294: 57-64.

32.Qing-Jun Yuan#, Zhi-Yong Zhang#, Lu-Qi Huang, Juan Hu, Lan-Ping Guo and Ai-Juan Shao. 2010. Impacts of modern cultivation on genetic diversity pattern of a medicinal plant, Scutellaria baicalensis (Lamilaceae). BMCGenetics, 11:29.

33.Zhi-Rong Zhang, Lai-Chun Luo, Ding Wuand Zhi-Yong Zhang*.2009. Two genetically distinct units of Sinomanglietia glauca (Magnoliaceae) detected by chloroplast PCR-SSCP. Journal of Systematics and Evolution 47:110-114.

34.Shuang Tian, Lai-Chun Luo, Song Ge and Zhi-YongZhang*. 2008. Clear population structure of Pinus kwangtungensis (Pinaceae) revealed by a plastid DNA fragment with a novel minisatellite. Annals of Botany 102:69-78.

35.Qing-Jun Yuan, Zhi-Yong Zhang*, Hua Peng* andSong Ge. 2008. Chloroplast phylogeography of Dipentodon (Dipentodontaceae) in southwest China and northernVietnam. Molecular Ecology,17:1054-1065.

36.Zhi-Yong Zhang, Xiao-Ming Zheng and Song Ge*. 2007. Population genetic structure of Vitex negundo (Verbenaceae) inThree-Gorge Area of the Yangtze River: the river barriers to seed dispersal in plants. Biochemical Systematics and Ecology, 35: 506-516.

37.Zhi-Yong Zhang, Yong-Yan Chen and De-Zhu Li*. 2005. Detection of lowgenetic variation in a critically endangered Chinese pine, Pinus squamatausing RAPD and ISSR markers. Biochemical Genetics, 43(5):239-249.

38.Zhi-Yong Zhang and De-Zhu Li*. 2004. Molecular phylogeny of section Parryaof Pinus based on matK gene sequence data. Journal of Integrative Plant Biology (formerly Acta BotanicaSinica), 46(2):171-179.

39.Zhi-Yong Zhang, Jun-Bo Yang and De-Zhu Li*.2003.Phylogenetic relationship of the extremely endangered species, Pinus squamata (Pinaceae) inferred from four sequences of the chloroplast genome and ITS of the nuclear genome. Journal of Integrative Plant Biology (formerly Acta Botanica Sinica), 45(5): 530-535.